CHECKLIST OF THE HORNWORTS AND LIVERWORTS OF ARKANSAS

Plantae
Anthocerotophyta
Anthocerotopsida
Anthocerotales
Anthocerotaceae Dum.
Anthoceros adscendens Lehin. and Lindenb.
Anthoceros ravenelii Aust.
Anthoceros punctatus L.
Anthoceros crispulus (Mont.) Douin
Notothylas orbi-cularis (Schwein.) Sull.
Phaeoceros laevis (L.) Prosk.
Phaeoceros laevis laevis
Anthoceros laevis L.
Phaeoceros laevis (L.) Prosk.
Phaeoceros laevis carolinianus (Michx.) Prosk.
Anthoceros carolinianus Michx.
Hepatophyta
Jungermanniopsida
Jungermanniales
Ptilidiineae
Tricholeaceae Nakai
Trichocholea tomentella (Ehrh.) Dum.
Lepidoziineae
Lepidoziaceae Limpr.
Bazzania trilobata (L.) S. Gray
Kurzia sylvatica (Evans) Grolle
Microlepidozia sylvatica (Evans) Joerg.
Calypogejaceae (K. Mull and S. Arnell)
Calypogeja fissa (L.) Raddi
Calypoqeja fissa neogaea Schust.
Calypoqeja muelleriama (Schiffn.) K. Mull.
Calypogeja sullivantii Aust.
Calypogeja arguta
var. sulhivantii (Aust. Frye and Clark)
Calypogeja trichomanis (L.) Corda
Cephaloziaceae Migula
Cephalozia biscuspidata (L.) Dum.
Cephalozia lunilifolia (Dum.) Dum.
Cephalozia media Lindb.
Adelanthaceae (Jeorg.) Grolle
Odontoschisma prostratum (Sw.) Trev.
Odontoschisma sphagni (Dicks.) Dum.
Cephaloziellaceae Douin
Cephaloziella divaricata (Sm.) Schiffn.
Cephaloziella divaricata var. divaricata
Cephaloziella byssacea (Roth) Warnst. ambiguum Nomen Nudum
Geocalycineae
Geocalycaceae Klinggr.
Geocalyx graveolams (Schrad.) Nees
Lophocoleaceae (Joerg.) V and Bergh.
Chiloscyphus pallescems (Ehrh. ex Hoffm.) Dum.
Chi-loscyphus pallescems var. fragilis (Roth) K. Mull.
Chiloscyphus polyamthos (L.) Corda
Lophoco lea bidentata (L.) Dum.
Lophocoleci cuspidata (Nees). Limpr.
Lophocoleci cuspidata var. cuspidata
Lophoco lea heterophylla (Schrad.) Dum.
Plagiochilaceae (Joerg.) K. Mull.
Plagiochila acanthophylla Gott.
Plagiochila acanthophylla japomica (Sande Lac.) H. Inoue
Plagiochila acanthophylla var. cihiligera (Schust.) H. Inoue
Plagiochila ludoviciciama Sull.
Plagiochila porelloides (Torrey ex Nees) Lindenb.
Plagiochila asplemioides (L.) Dum.
Plagiochila undata Sull.
Plagiochila undata undata
Jungermanniineae
Jungermanniaceae
Jamesomiella autumnalis (DC.) Steph.
Jungermammia hyalina Lyell
Plectoco leahyailina (Lyell) Mitt.
Jungermannia leicintha Grolle
Jungermannia lanceolata Auct.
Gynnomitriaceae Klinggr.
Marsupella sphacelata (Gieseke) Dum.
Marsupella sulhivantii (DeNot) Evans
Scapaniaceae Migula
Diplophyllum apiculatum (Evans) Steph.
Diplophyllum apiculatum var. apiculatum
Scapamia irriguci (Nees) Gott. et al.
Scapamia memorosa (L.) Dum.
Scapamia undulata (L.) Dum.
Scapamia undulata var. undulata
Radulineae
Radulaceae (Dum.) K. Mull.
Raddula mollis Lindenb. and Gott.
Radula andicola Steph.
Radula oboomica Sull.
Porellineae
Porellaceae
Porella pinnata L.
Porella platyphylla (L.) Pfeiff.
Porella platyphylloides (Schwein.) Lindb.
Porella platyphylla var. platyphylloideci (Schwein.) Frye and Clark
Jubulaceae Klinggr.
Frullamia brittoniae Evans
Frullania donnellii Aust.
Frullania eboracensis Gott.
Frullania virginica Gott.
Frullania inflata Gott.
Frullania kunzei Lehin. and Lindenb.
Frullania plana Sull.
Frullania riparia Hampe ex Lehm
Frullania squarrosa (Reinw. et al.) Dum.
Frullania tamarisci (L.) Dum.
Frullania tamarisci asagrayana (Mont.) Hatt.
Frullania asagraycina Mont.
Jubula pennsylvanica (Steph.) Evans
Leujeuneaceae Gas. Gil.
Cololejeunea biddle comae (Aust.) Evans
Lejeunea lamacerima (Steph.) Schiffn.
Leucolejeumeia clypeata (Schwein.) Evans
Archilejeuneia clypeata (Schwein.) Schiffn.
Phragrmicoma clypeata Sulliv.
Leucolejeumea unciloba (Lindenb.) Evans
Microlejeunca lactevirens (Nees and Mont.) Evans
Codoniineae
Codoniaceae Klinggr.
Fossombronia brasihiensis Steph.
Possombronia foveolata Lindb.
Fossombromia womdraczekui (Corda) Dum.
Petalophyllum ralfsi (Wils.) Nees and Gott.
Petalophyllum lamellatum (Nees) Lindb.
Metzgeriineae
Pallaviciniaceae Migula
Pallavicimia lyellii (Hook.) Carruth.
Pelliaceae Klinggr.
Pellia endiviifolia (Dicks.) Dum.
Pellia epiphylla (L.) Corda
Pelliia nessiana (Gott.) Limpr.
Metzgeriaceae Klinggr.
Metzgena conjugata Lindb.
Metzgeria crassipilis Lindb.
Metzgeria furcata (L.) Dum.
Aneuraceae Klinggr.
Aneura pinguis (L.) Dumb.
Riccardia pinguis (L.) S. Gray
Riccardia latifrons Lindb.
Riccardia multifida (L.) S. Gray
Riccardia palmata (Hedw.) Carruth.
Ameura palmata Nees.
Marchantiopsida
Sphaerocarpiidae
Sphaerocarpales
Sphaerocarpaceae
Sphaerocarpos michelii Bell.
Sphaerocarpos texanus Aust.
Marchantiidae
Marchantiales
Marchantiineae
Aytoniaceae Cavers
Asterella elegans (Spreng.) Trev.
Asterella elegans echinella (Gott.) del Rosario
Asterella echinella (Gott.) Underw.
Asterella tenella (L.) P. Beauv.
Mannia califormica (Gott.) Wheeler
Mannia fragrans (Balbis) Frye and Clark
Mannia triandra (Scop.) Grolle
Asterella ludwigii (Schwaegr.) Underw.
Mannia ruprestris (Nees) Frye and Clark
Reboulia hemisphacrica (L.) Raddi
Asterella hemisphenica Beauv.
Conocephalaceae K. Mull.
Conocephalum conicum (L.) Lindb.
Conocephalus conicus Dum.
Lunulariaceae Klinggr.
Lunulania cruciata (L.) Dum.
Marchantiaceae (Bisch.) Endl.
Dumortiera hirsuta (Sw.) Nees
Dumortiera hirsuta var. hirsuta (SW.) Nees
Marchantia domingensis Lehm. and Lindenb.
Marchantia polymorpha L.
Preissia quadrata (Scop.) Nees
Ricciineae
Ricciaceae Reichenb.
Riccia austini Steph.
Riccia beyrichiana Hampe ex Lehm.
Riccia campbellaina M.A. Howe
Riccia cavernosa Hoffm.
Riccia crystallina Auct.
Riccia curtisii (Aust.) James
Riccia dictyospora M.A. Howe
Riccia fluitans L.
Riccia frostii Aust.
Riccia hirta (Aust.) Underw.
Riccia macallisteri M.A. Howe
Riccia membranaceci Cott. and Lindenb.
Riccia tenuis Aust.
Riccia ozarkiana McGregor
Riccia sorocarpa Bisch.
Riccia sullivanti Aust.
Riccia fluitans var. sullivantii Aust.
Ricciocarpos natans (L.) Corda

Literature cited:
Branner, J.C., and F.V. Coville. 1891. A list of the plants of Arkansas, Annual Report of the Geological Survey of Arkansas for 1888. Ark. Geol. Surv. 4:155-242. Press Printing Co., Little Rock, Arkansas.
Coulter, S.M. 1904. An ecological comparison of some typical swamp areas. 15th Ann. Rep. Missouri Bot. Gard. :38-40. St. Louis, Mo.
Evans, A.W. 1902. The Lejeuneae of the United States and Canada. Memoirs Torrey Bat. Clb. VIII:113-194.
Evans, A.W. 1903. Odontoschi-sma macouni-i and its North American allies. Bot. Gaz. 36:321-348.
Evans, A.W. 1907. Notes on New England Hepaticae 5. Rhodora 9:56-60; 65-73.
Evans, A.W. 1917. Notes on New England Hepaticae 14. Rhodora 19:263-272.
Evans, A.W. 1920. The North American species of Asterella. Contr. U.S. Nat. Herb. 20:247-312. Evans, A.W. 1923. Family Rebouliaceae and Family Marchantiaceae in North American Flora. Vol. 14, Part 1.
Evans, A.W., and G. E. Nichols. 1908. The bryophytes of Connecticut. Conn. State Geol. Nat. Hist. Surv. Bull. 11:1-203. Hartford.
Fulford, Margaret. 1936. Additions to the list of Hepaticae of Kentucky 1. Bryologist 39:119-122.
Howe, M.A. 1897. The North American species of Porella. Bull. Torr. Bot. Club 24:512-528.
Jacobs, D.L. 1951. Ricciaceae of Arkansas. The Bryologist 54:274-278.
Kurz, H. and T.M. Little. 1933. Liverworts of north and central Florida. Bull. Florida State College for Women 26:1-41.
Lesquereux, Leo. 1860. Botanical and paleontological report on the Geological State Survey of Arkansas. Geol. Surv. of Ark. 2:295-399. Philadelphia.
Lorenz, Annie. 191&. Notes on Padula obconi-ca Sull. Bryologist 21:56-59.
McGregor, Ronald L. 1960. A new species of Riccia from the Ozarks. The Bryologist 63:30-31.
Moore, Jewel E. 1965. A study of the vegetation of Petit Jean Mountain in central Arkansas. Castanea 30:1-37.
Moore, Jewel E., and Steve L. Tiimne. 1983. Bryophyte and lichen communities within Hot Springs National Park, Arkansas. In prep. for Ark. Acad. Sci. Proc.
Redfearn, Paul L., Jr. 1964. Bryophytes of Arkansas I. species of “Lost Valley” and adjacent regions. The Bryologist 67:196-201.
Redfearn, Paul L., Jr. 1966. Bryophytes of the Interior Highlands of North America. XI. Additions to the Flora. The Bryologist 69:504-508.
Redfearn, Paul L., Jr. 1968. Bryophytes from the Interior Highlands. XIII. Additions to the Flora. The Bryologist 71:356-357.
Redfearn, Paul L., Jr. 1970. Bryophytes of the Interior Highlands XV. Additions to the Flora. The Bryologist 73:716-717.
Redfearn, Paul L., Jr. 1979. Bryophytes of the Interior Highlands of North America. XVI. Species of the Ozark National Forest, Sylamore Division, and adjacent areas in Stone County, Arkansas. The Bryologist 82:483-486.
Schuster, Rudolf M. 1966. The Hepaticae and Anthocerotae of North America, east of the hundredth meridian. Vol. 1. Columbia University Press. New York.
Stephani, F. 1899. Species Hepaticorum. Bull. Herb. Boiss. 7:109.
Stotler, Raymond, and Barbara Crandall-Stotler. 1977. A checklist of the liverworts and hornworts of North America. The Bryologist 80:405-428.
Underwood, L.M. 1895. Notes on our Hepaticae III. The distribution of North American Marchantiaceae. Bot. Gaz. 20:59-71.
Wittlake, Eugene B. 1950A. Preliminary report on the Arkansas bryophytes. Ark. Acad. Sci. Proc. 3:35-37.
Wittlake, Eugene B. 1950B. The bryophytes of Spy Rock Hollow. Ark. Acad. Sci. Proc. 3:39-40.
Wittlake, Eugene B. 1951. A new record for Petalophyllum lamellatum (Nees) Lindb. The Bryologist 54:76.
Wittlake, Eugene B. 1954. The Hepaticae of Arkansas I. The Bryologist 57:7-18.
Wittlake, Eugene B. 1972. 1972. Bryophytes of Strawberry River Basin. Appendix C of Environmental Inventory, Strawberry River Basin. Prepared by Ark. State Univ. for U.S. Army Corps District, Corps of Engineers, Little Rock, Ark.

Source:
Moore, Jewel E. and Eugene B. Wittlake. 1983. Systematic checklist of the Arkansas Hornworts and Liverworts. Original contribution. University of Central Arkansas, Conway. Arkansas State University, State University, Arkansas

This checklist is to be cited as:
Moore, Jewel E. and Eugene B. Wittlake. 1983. Checklist of the Hornworts and Liverworts of Arkansas.
Ark. Acad. of Science, Ark. Biota Survey Checklist No. 36. 7pp.

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